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Fire ants with Sb and SB haplotypes only retain the SB copy, whereas both Si OR88 and Si OR89 genes lie outside the nine-exon clade, which has significantly expanded due to the duplication of OR genes ( Si OR463, Si OR88, and Si OR89 have been linked to the social structure of RIFA colonies, including their ability to recognize queens (Wicher and Marion-Poll, 2018
, Sakarcan, A
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